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He added: "I would continue to suggest that if Gareth was not killed as a result of a sex crime or sex game gone wrong - and nobody has presented any serious evidence of this - then this must be the work of a hostile intelligence service.
Prof Glees, 62, said Mr Williams may have been blackmailed into passing on secrets to a hostile intelligence service which wanted to leave no trace of any links back to its organisation.
Intelligence Online reported on 15 October that Aghazadeh, who is also a member of the influential Council of Expediency, was dismissed as head of the nuclear agency for allowing Amiri to move out of the Greater Tehran area without security surveillance, a measure intended to prevent defections or kidnappings by hostile intelligence services such as the CIA or the Mossad.